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A version of this paper originally appeared in INSECTA MUNDI 4(1):247-254 (1986)

Keep in mind that it has not been updated since then.

A CATALOG OF THE MAYFLIES (EPHEMEROPTERA) OF HONG KONG

Michael D. Hubbard
Laboratory of Aquatic Entomology
Florida A&M University
Tallahassee, Florida 32307-4100 USA

This paper is a catalog of the Ephemeroptera known from the British Crown Colony of Hong Kong. It includes all known references for each species, along with an indication of the type localities and depositions of type specimens. For each reference the stage (male, female, or nymph) is indicated if relevant. Keys which include the species also are indicated. The recorded distribution for each species is listed. For each genus are given the reference to the original description, information on the type species, and any synonyms.

Abbreviations of places of type deposition are as follows: (BMNH), British Museum (Natural History), London; (Cornell), Cornell University, Ithaca, New York; (DEI), Deutsches Entomologisches Institut, Eberswalde; (Hamburg), Zoologisches Staatsinstitut und Zoologisches Museum, Hamburg; (Hsu), personal collection of Y.-C. Hsu [the late Prof. Hsu indicated (personal communication) that because of events in China over the past few decades these specimens should be considered destroyed or lost]; (Utah), University of Utah, Salt Lake City.

Although the taxonomic knowledge of the Hong Kong mayflies is still somewhat sketchy, there are substantial ecological studies ongoing at this time. Therefore, in order to facilitate future studies and comparisons of data, I have included in this catalog the "morphospecies" of Dudgeon (1982a, b, 1983, 1984a, b). The reader should be aware that many of the taxonomic assignments of these morphospecies are preliminary and actual placement will have to await future taxonomic studies.

This paper is one in a continuing series of catalogs of the Ephemeroptera (cf. Hubbard & Peters 1978, Hubbard & Pescador 1978, Hubbard & Savage 1981, Hubbard 1982a,b).

I thank Prof. David Dudgeon of the University of Hong Kong for his helpful comments in the preparation of this manuscript.

This paper has been partially supported by a grant from the Cooperative States Research Service, U. S. Department of Agriculture, to the Agriculture Research Programs at Florida A&M University.

Order Ephemeroptera

Family Baetidae Leach, 1815

Subfamily Baetinae Leach, 1815

Genus Baetis Leach, 1815

  • Baetis Leach, 1815:137.
    • Type species: Ephemera fuscata Linnaeus. Subsequent designation by Intern. Comm. Zool. Nomencl., 1966:209.
  • Brachyphlebia Westwood, 1840:25.
    • Type species: Ephemera fuscata Linnaeus (misidentified as Ephemera bioculata) [cf. Intern. Comm. Zool. Nomencl., 1966:209]. Monotypic.
  • Acentrella Bengtsson, 1912:110.
    • Type species: Acentrella laponica Bengtsson. Monotypic.
  • Heterocloeon McDunnough, 1925:175.
    • Type species: Centroptilum curiosum McDunnough. Original designation.

Baetis sp. T1

  • Baetis T1 Dudgeon, 1982:14.
  • Baetis T1 Dudgeon, 1982:225.
  • Baetis T1 Dudgeon, 1983:441.
  • Baetis T1 Dudgeon, 1984:73.
  • Baetis T1 Dudgeon, 1984:210.
    • Distribution: Hong Kong.

Baetis sp. T2

  • Baetis T2 Dudgeon, 1982:14.
  • Baetis T2 Dudgeon, 1982:225.
  • Baetis T2 Dudgeon, 1983:441.
  • Baetis T2 Dudgeon, 1984:73.
  • Baetis T2 Dudgeon, 1984:210.
    • Distribution: Hong Kong.

Baetis sp. T3

  • Baetis T3 Dudgeon, 1982:14.
  • Baetis T3 Dudgeon, 1982:225.
  • Baetis T3 Dudgeon, 1983:441.
  • Baetis T3 Dudgeon, 1984:73.
  • Baetis T3 Dudgeon, 1984:210.
    • Distribution: Hong Kong.

Baetis sp. T4

  • Baetis T4 Dudgeon, 1982:225.
  • Baetis T4 Dudgeon, 1983:441.
  • Baetis T4 Dudgeon, 1984:73.
  • Baetis T4 Dudgeon, 1984:210.
    • Distribution: Hong Kong.

Baetis sp. T5

  • Baetis T5 Dudgeon, 1984:73.
  • Baetis T5 Dudgeon, 1984:210.
    • Distribution: Hong Kong.

Baetis sp. L5

  • Baetis L5 Dudgeon, 1984:73.
  • Baetis L5 Dudgeon, 1984:210.
    • Distribution: Hong Kong.

Baetis sp. L6

  • Baetis L6 Dudgeon, 1984:73.
  • Baetis L6 Dudgeon, 1984:210.
    • Distribution: Hong Kong.

Baetis sp. L7

  • Baetis L7 Dudgeon, 1984:73.
  • Baetis L7 Dudgeon, 1984:210.
    • Distribution: Hong Kong.

Baetis sp. L8

  • Baetis L8 Dudgeon, 1984:73.
  • Baetis L8 Dudgeon, 1984:210.
    • Distribution: Hong Kong.

Baetis sp. cf. T3

  • Baetis cf. T3 Dudgeon, 1984:73.
  • Baetis cf. T3 Dudgeon, 1984:210.
    • Distribution: Hong Kong.

Subfamily Cloeoninae Newman, 1853

Genus Centroptilum Eaton, 1869

  • Centroptilum Eaton, 1869:132.
    • Type species: Ephemera luteola Müller. Original designation.

Centroptilum sp. L1

  • Centroptilum L1 Dudgeon, 1984:73.
  • Centroptilum L1 Dudgeon, 1984:210.
    • Distribution: Hong Kong.

Genus Cloeon Leach, 1815

  • Cloeon Leach, 1815:137.
    • Type species: Ephemera diptera Linnaeus as Cloeon pallidus Leach. Monotypy.
  • Cloe Burmeister, 1839:797. [unjustified emendation of Cloeon Leach].
    • Type species: Ephemera diptera Linnaeus. Objective synonymy.
  • Cloeopsis Eaton, 1866:146.
    • Type species: Ephemera diptera Linnaeus. Monotypy.
  • Austrocloeon Barnard, 1932:217. [nomen nudum].
  • Neoc1oeon Traver, 1932:365.
    • Type species: Neocloeon alamance Traver. Original designation.
  • Austrocloeon Barnard, 1940:616.
    • Type species: Cloeon africanum Esben Petersen. Original designation.

Cloeon harveyi Kimmins

  • Procloeon harveyi Kimmins, 1947:94, fig. 2, 6, 10. (male, female). Type: female, Calcutta, Bengal (BMNH).
  • Procloeon harveyi - Gillies, 1949:176.
  • Procloeon harveyi - Kimmins, 1971:314.
  • Procloeon harveyi - Hubbard & Peters, 1978:11.
  • Procloeon harveyi - Hubbard & Srivastava, 1984:3.
  • Cloeon harveyi - Müller-Liebenau & Hubbard, 1986:538.
    • Distribution: Hong Kong, Malaysia, India.

Genus Pseudocloeon Klapálek, 1905

  • Pseudocloeon Klapá1ek, 1905:105.
    • Type species: Pseudocloeon kraepelini Klapálek. Monotypic.
  • Baetiella Uéno, 1931:220.
    • Type species: Acentrella japonica Imanishi. Original designation.

Pseudocloeon sp. T1

  • Pseudocloeon T1 Dudgeon, 1982:14.
  • Pseudocloeon T1 Dudgeon, 1982:225.
  • Pseudocloeon T1 Dudgeon, 1983:441.
  • Pseudocloeon T1 Dudgeon, 1984:73.
  • Pseudocloeon T1 Dudgeon, 1984:210.
    • Distribution: Hong Kong.

Pseudocloeon sp. T2

  • Pseudocloeon T2 Dudgeon, 1982:14.
  • Pseudocloeon T2 Dudgeon, 1982:225.
  • Pseudocloeon T2 Dudgeon, 1983:441.
  • Pseudocloeon T2 Dudgeon, 1984:73.
  • Pseudocloeon T2 Dudgeon, 1984:210.
    • Distribution: Hong Kong.

Pseudocloeon sp. L3

  • Pseudocloeon L3 Dudgeon, 1984:73.
  • Pseudocloeon L3 Dudgeon, 1984:210.
    • Distribution: Hong Kong.

Pseudocloeon sp.

  • Baetella [sic] T1 Dudgeon, 1983:441.
    • Distribution: Hong Kong.

Family Caenidae Newman, 1853

Genus Caenis Stephens, 1835

  • Caenis Stephens, 1835:60.
    • Type species: Caenis macrura Stephens. Subsequent designation by Westwood, 1840:47.
  • Oxycypha Burmeister, 1839:796.
    • Type species: Oxycypha luctuosa Burmeister. Subsequent designation by Jacob, 1974:96.
  • Ordella Campion, 1923:518.
    • Type species: Caenis macrura Stephens. Original designation.

Caenis sp. T1

  • Caenis T1 Dudgeon, 1982:14.
  • Caenis T1 Dudgeon, 1982:225.
  • Caenis T1 Dudgeon, 1984:73.
  • Caenis T1 Dudgeon, 1984:210.
    • Distribution: Hong Kong.

Caenis sp. T2

  • Caenis T2 Dudgeon, 1983:441.
  • Caenis T2 Dudgeon, 1984:73.
  • Caenis T2 Dudgeon, 1984:210.
    • Distribution: Hong Kong.

Caenis sp. L2

  • Caenis L2 Dudgeon, 1984:73.
  • Caenis L2 Dudgeon, 1984:210.
    • Distribution: Hong Kong.

Caenis sp. L3

  • Caenis L3 Dudgeon, 1984:73.
  • Caenis L3 Dudgeon, 1984:210.
    • Distribution: Hong Kong.

Family Ephemeridae Latreille, 1810

Genus Ephemera Linnaeus, 1758

  • Ephemera Linnaeus, 1758:546.
    • Type species: Ephemera vulgata Linnaeus. Subsequent designation by Westwood, 1836:439.
  • Nirvius Navás, 1922:56.
    • Type species: Nirvius punctatus Navás. Original designation.

Subgenus Ephemera Linnaeus, 1758

Ephemera (Ephemera) pulcherrima Eaton

  • Ephemera pulcherrima Eaton, 1892:185. (female). Type: female, Tenasserim Valley, Burma (BMNH).
  • Ephemera pulcherrima - Ulmer, 1926:86, fig. 80-81. (male, key).
  • Ephemera pulcherrima - Lestage, 1927:99. (key).
  • Ephemera pulcherrima - Wu, 1935:248.
  • Ephemera pulcherrima - Hsu, 1936:233, fig.1.(female).
  • Ephemera pulcherrima - Ulmer, 1936:204.
  • Ephemera pulcherrima - Hafiz, 1937:368.
  • Ephemera pulcherrima - Hsu, 1937:433, fig. 25-26. (male, female, key).
  • Ephemera pulcherrima - Kimmins, 1960:308.
  • Ephemera pulcherrima - Kimmins, 1971:319.
  • Ephemera pulcherrima - Hubbard & Peters, 1978:17.
    • Distribution: Hong Kong, China, Burma, India.

Ephemera (Ephemera) serica Eaton

  • Ephemera serica Eaton, 1871:75, p1. 4 fig. 12-12a. (male, female). Type: N. China (BMNH).
  • Ephemera serica - McLachlan, 1875:169.
  • Ephemera serica - Eaton, 1883:73. (male, female).
  • Ephemera serica - Ulmer, 1926:89, fig. 82-83. (male, female, key).
  • Ephemera serica - Lestage, 1927:99. (key).
  • Ephemera serica - Navás, 1930:15.
  • Ephemera serica - Navás, 1930:138.
  • Ephemera serica - Wu, 1935:248.
  • Ephemera serica - Ulmer, 1936:206.
  • Ephemera serica - Hsu, 1937:438. (male, female, key).
  • Ephemera serica - Kimmins, 1960:309, fig. 53. (male). [designation of lectotype].
  • Ephemera serica - Kimmins, 1971:320.
    • Distribution: Hong Kong, China, Vietnam.

Ephemera (subgenus incertus)

Ephemera sp. T1

  • Ephemera T1 Dudgeon, 1982:225.
    • Distribution: Hong Kong.

Ephemera sp. T2

  • Ephemera T2 Dudgeon, 1983:441.
    • Distribution: Hong Kong.

Ephemera sp.

  • Ephemera sp. Dudgeon, 1984:73.
  • Ephemera sp. Dudgeon, 1984:210.
    • Distribution: Hong Kong.

Family Ephemerellidae Klapálek, 1909

Subfamily Ephemerellinae Klapálek, 1909

Genus Ephemerella Walsh, 1863

  • Ephemerella Walsh, 1863:377.
    • Type species: Ephemerella excrucians Walsh. Subsequent designation by Eaton, 1868:87.

Ephemerella sp. T1

  • Ephemerella T1 Dudgeon, 1982:14.
  • Ephemerella T1 Dudgeon, 1982:225.
  • Ephemerella T1 Dudgeon, 1983:441.
  • Ephemerella T1 Dudgeon, 1984:73.
  • Ephemerella T1 Dudgeon, 1984:210.
    • Distribution: Hong Kong.

Ephemerella sp. T2

  • Ephemerella T2 Dudgeon, 1982:14.
  • Ephemerella T2 Dudgeon, 1982:225.
  • Ephemerella T2 Dudgeon, 1984:73.
  • Ephemerella T2 Dudgeon, 1984:210.
    • Distribution: Hong Kong.

Ephemerella sp. L3

  • Ephemerella L3 Dudgeon, 1984:73.
  • Ephemerella L3 Dudgeon, 1984:210.
    • Distribution: Hong Kong.

Family Heptageniidae Needham in Needham & Betten, 1901

Subfamily Heptageniinae Needham in Needham & Betten, 1901

Genus Compsoneuria Eaton, 1881

  • Compsoneuria Eaton, 1881:23.
    • Type species: Compsoneuria spectabilis Eaton. Original designation.
  • Cosmetogenia Eaton, 1883:18, p1. 23.
    • Type species: Compsoneuria spectabilis Eaton. Monotypy.

Compsoneuria sp. T1

  • Ecdyonurus T1 Dudgeon, 1982:14.
  • Ecdyonurus T1 Dudgeon, 1982:225.
  • Ecdyonurus T1 Dudgeon, 1983:441.
  • Ecdyonurus T1 Dudgeon, 1984:73.
  • Compsoneuria T1 Dudgeon, 1984:210.
    • Distribution: Hong Kong.

Compsoneuria sp. T2

  • Ecdyonurus T2 Dudgeon, 1982:225.
  • Ecdyonurus T2 Dudgeon, 1984:73.
  • Compsoneuria T2 Dudgeon, 1984:210.
    • Distribution: Hong Kong.

Genus Ecdyonurus Eaton, 1868

  • Ecdyonurus Eaton, 1868:142.
    • Type species: Ephemera venosa Fabricius. Original designation.
  • Ecdyurus Eaton, 1868:90.
    • Type species: Ephemera venosa Fabricius. Objective synonymy. [unjustifed emendation of Ecdyonurus].
  • Paracinygmula Bajkova, 1975:54.
    • Type species: Paracinygmula zhilzovae Bajkova. Original designation.

Ecdyonurus herklotsi Hsu, 1936

  • Ecdyonurus herklotsi Hsu:233, fig. 2-4. (male). Type: male, Pokfulum [Pokfulam], Hong Kong (Hsu).
    • Distribution: Hong Kong.

Genus Epeorus Eaton, 1881

  • Epeorus Eaton, 1881:26.
    • Type species: Epeorus torrentium Eaton. Original designation.

Subgenus Iron Eaton, 1883

  • Iron Eaton, 1883: p1. 24.
    • Type species: Epeorus longimanus Eaton. Monotypy.

Epeorus (Iron) sp.

  • Iron sp. Dudgeon, 1984:73.
  • Iron sp. Dudgeon, 1984:210.
    • Distribution: Hong Kong.

Epeorus (subgenus incertus)

Epeorus sp. T1

  • Epeorus T1 Dudgeon, 1982:14
  • Epeorus T1 Dudgeon, 1982:225.
  • Epeorus T1 Dudgeon, 1983:441.
  • Epeorus T1 Dudgeon, 1984:73.
  • Epeorus T1 Dudgeon, 1984:210.
    • Distribution: Hong Kong.

Epeorus sp. T2

  • Epeorus T2 Dudgeon, 1984:73.
  • Epeorus T2 Dudgeon, 1984:210.
    • Distribution: Hong Kong.

Genus Heptagenia Walsh, 1863

  • Heptagenia Walsh, 1863:197.
    • Type species: Palingenia flavescens Walsh. Subsequent designation by Eaton, 1868, Entomol. Mon. Mag. 5:90.
  • Kageronia Matsumura, 1931:1479.
    • Type species: Kageronia suzukiella Matsumura. Monotypic.

Heptagenia ngi Hsu, 1936

  • Heptagenia ngi Hsu, 1936:235, fig. 5-7. (male). Type: male, Shaukiwan, Hong Kong (Hsu).
    • Distribution: Hong Kong.

Genus Paegniodes Eaton, 1881

  • Paegniodes Eaton, 1881:23.
    • Type species: Heptagenia cupulata Eaton. Original designation.

Paegniodes cupulatus (Eaton, 1871)

  • Heptagenia cupulata Eaton, 1871:138, p1. 6 fig.
    • 14-14a. (male, female). Type: Northern China (BMNH).
  • Paegniodes cupulatus - Eaton, 1881:23.
  • Heptagenia cupulata - Palmén, 1884:74.
  • Paegniodes cupulatus - Eaton, 1885:261, p1. 23 fig. 4, p1. 24 fig. 4. (male, female).
  • Paegniodes cupulatus - Ulmer, 1926:107, figs. 98-102. (male, female).
  • Paegniodes cupulatus - Hsu, 1932:39.
  • Paegniodes fukienensis - Hsu, 1932:39, fig. 1-3. (male). Type: male, Ling Sioh, Fuken Prov. China (Cornell).
  • Paegniodes cupulatus - Wu, 1935:252.
  • Paegniodes fukienensis - Wu, 1935:252.
  • Paegniodes cupulata - Hsu, 1936:236, fig. 8-13. (male, female).
  • Paegniodes cupulatus - Ulmer, 1936:211. (male).
  • Paegniodes cupulatus - Kimmins, 1960:283. [designation of lectotype].
  • Paegniodes cupulatus - Uéno, 1969:228, fig. 26-36. (male).
  • Heptagenia cupulata - Kimmins, 1971:312.
  • Paegniodes cupulatus - Tshernova, 1976:343, fig. 41. (nymph).
  • Paegniodes fukiensis [sic] - Tshernova, 1976:343.
    • Distribution: Hong Kong, China, Tibet.

Genus Thalerosphyrus Eaton, 1881

  • Thalerosphyrus Eaton, 1881:22.
    • Type species: Baetis determinatus Walker. Original designation.

Thalerosphyrus sp. T1

  • Thalerosphyrus T1 Dudgeon, 1982:14.
  • Thalerosphyrus T1 Dudgeon, 1983:441.
  • Thalerosphyrus T1 Dudgeon, 1984:73.
  • Thalerosphyrus T1 Dudgeon, 1984:210.
    • Distribution: Hong Kong.

Family Leptophlebiidae Banks, 1900

Subfamily Atalophlebiinae Peters, 1980

Genus Choroterpes Eaton, 1881

  • Choroterpes Eaton, 1881:194.
    • Type species: Choroterpes lusitanica Eaton. Original designation.

Subgenus Euthraulus Barnard, 1932

  • Euthraulus Barnard, 1932:249.
    • Type species: Euthraulus elegans Barnard. Monotypy.
  • Thraululus Ulmer, 1939:499.
    • Type species: Thraulus marginatus Ulmer. Original designation.

Choroterpes (Euthraulus) sp.

  • Choroterpes (Euthraulus) sp. Dudgeon, 1984:73.
  • Choroterpes (Euthraulus) sp. Dudgeon, 1984:210.
    • Distribution: Hong Kong.

Choroterpes (subgenus incertus)

Choroterpes sp. T1

  • Choroterpes T1 Dudgeon, 1982:14.
  • Choroterpes T1 Dudgeon, 1983:441.
    • Distribution: Hong Kong.

Genus Cryptopenella Gillies, 1951

  • Cryptopenella Gillies, 1951:125.
    • Type species: Cryptopenella facialis Gillies. Original designation.

Cryptopenella facialis Gillies, 1951

  • Cryptopenella facialis Gillies, 1951:127, fig. 11-12, 14. (male, female, nymph). Type: River Shing Man (Shing Mum River], near Kowloon, Hong Kong (BMNH).
  • Cryptopenella facialis - Peters & Edmunds, 1970:202, fig. 46-47, 93-94, 122, 145, 336-345. (male, female, nymph).
  • Cryptopenella facialis - Kimmins, 1971:313.
    • Distribution: Hong Kong.

Genus Habrophlebiodes Ulmer, 1920

  • Habrophlebiodes Ulmer, 1920:39.
    • Type species: Habrophlebia americana Banks. Original designation.

Habrophlebiodes gilliesi Peters, 1963

  • Habrophlebiodes gilliesi Peters, 1963:41, fig. 1-11. (male, female). Type: male, Fan Ling near Kowloon, Hong Kong (Utah).
  • Habrophlebiodes gilliesi - Peters & Edmunds, 1970:187, fig. 19-21, 77, 112, 135, 323. (male, female).
  • Habrophlebiodes gilliesi - Tsui & Peters, 1970:89, fig. 1-13. (nymph).
  • Habrophlebiodes gilliesi - Dudgeon, 1982:225.
    • Distribution: Hong Kong.

Genus Isca Gillies, 1951

  • Isca Gillies, 1951:127.
    • Type species: Isca purpurea Gillies. Original designation.

Subgenus Isca Gillies, 1951

Isca (Isca) purpurea Gillies, 1951

  • Isca purpurea Gillies, 1951:128, fig. 15-22. (male, female). Types: male, female, Hong Kong, 15-20 mi inland from Kowloon (BMNH).
  • Isca (Isca) purpurea - Peters & Edmunds, 1970:215, fig. 71, 106, 330, 350-357. (male, nymph).
  • Isca purpurea - Kimmins, 1971:319.
  • Isca (Isca) purpurea - Hubbard & Peters, 1978:24.
    • Distribution: Hong Kong, India.

Isca (subgenus incertus)

Isca sp. T1

  • Isca T1 Dudgeon, 1982:225.
  • Isca T1 Dudgeon, 1983:441.
    • Distribution: Hong Kong.

Isca sp.

  • Isca sp. Dudgeon, 1984:73.
  • Isca sp. Dudgeon, 1984:210.
    • Distribution: Hong Kong.

Genus Thraulus Eaton, 1881

  • Thraulus Eaton, 1881:195.
    • Type species: Thraulus bellus Eaton. Original designation.
  • Masharikella Peters, Gillies & Edmunds, 1964:118.
    • Type species: Hagenulus fasciatus Kimmins. Original designation.

Thraulus sp. T1

  • Thraulus T1 Dudgeon, 1982:225.
  • Thraulus T1 Dudgeon, 1983:441.
    • Distribution: Hong Kong.

LITERATURE CITED

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Eaton, A. E. 1866. Notes on some species of the orthopterous genus Cloeon, Leach (as limited by M. Pictet). Annals and Magazine of Natural History (3)18:145-148.

Eaton, A. E. 1868a. Remarks upon the homologies of the ovipositor. Transactions of the Entomological Society 1868:141-144.

Eaton, A. E. 1868b. An outline of a rearrangement of the genera of Ephemeridae. Entomologist's Monthly Magazine 5:82-91.

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Eaton, A. E. 1871. A monograph on the Ephemeridae. Transactions of the Entomological Society 1871:1-164, 6 p1.

Eaton, A. E. 1881a. An announcement of new genera of the Ephemeridae. Entomologist's Monthly Magazine 17:191-197.

Eaton, A. E. 1881b. An announcement of new genera of the Ephemeridae. Entomologist's Monthly Magazine 18:21-27.

Eaton, A. E. 1883-1885. A revisional monograph of recent Ephemeridae or mayflies. Transactions of the Linnean Society of London, Second Series, Zoology 3:1-352, 65 p1.

Eaton, A. E. 1892. New species of Ephemeridae from the Tenasserim Valley. Transactions of the Entomological Society of London 1892:185-190.

Gillies, M. T. 1949. Notes on some Ephemeroptera Baëtidae from India and South-East Asia. Transactions of the Royal Entomological Society London 100:161-177.

Gillies, M. T. 1951. Further notes on Ephemeroptera from India and South East Asia. Proceedings of the Royal Entomological Society of London (B)20:121-130.

Hafiz, H. A. 1937. The Indian Ephemeroptera (mayflies) of the sub-order Ephemeroidea. Records of the Indian Museum 39:351-370, p1. 4-5.

Hsu, Y.-C. 1932. Two new species of may-flies from China (Order Ephemerida). Peking Natural History Bulletin 6:39-41, 1 p1.

Hsu, Y.-C. 1936. Mayflies of Hong Kong with descriptions of two new species (Ephemeroptera). Hong Kong Naturalist 7:233-238.

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